Introduction to the Muscular System

The muscular system is composed of specialized cells called muscle fibers. Their predominant function is contractibility. Muscles, attached to bones or internal organs and blood vessels, are responsible for movement. Nearly all movement in the body is the result of muscle contraction. Exceptions to this are the action of [glossary term:]cilia, the [glossary term:]flagellum on sperm cells, and [glossary term:]amoeboid movement of some white blood cells.

The integrated action of joints, bones, and skeletal muscles produces obvious movements such as walking and running. Skeletal muscles also produce more subtle movements that result in various facial expressions, eye movements, and respiration.

In addition to movement, muscle contraction also fulfills some other important functions in the body, such as posture, joint stability, and heat production. Posture, such as sitting and standing, is maintained as a result of muscle contraction. The skeletal muscles are continually making fine adjustments that hold the body in stationary positions. The tendons of many muscles extend over joints and in this way contribute to joint stability. This is particularly evident in the knee and shoulder joints, where muscle tendons are a major factor in stabilizing the joint. Heat production, to maintain body temperature, is an important by-product of muscle metabolism. Nearly 85 percent of the heat produced in the body is the result of muscle contraction.

Structure of Skeletal Muscle

A whole skeletal muscle is considered an organ of the muscular system. Each organ or muscle consists of skeletal muscle tissue, connective tissue, nerve tissue, and blood or vascular tissue.

Skeletal muscles vary considerably in size, shape, and arrangement of fibers. They range from extremely tiny strands such as the stapedium muscle of the middle ear to large masses such as the muscles of the thigh. Some skeletal muscles are broad in shape and some narrow. In some muscles the fibers are parallel to the long axis of the muscle; in some they converge to a narrow attachment; and in some they are oblique.

Each skeletal muscle fiber is a single cylindrical muscle cell. An individual skeletal muscle may be made up of hundreds, or even thousands, of muscle fibers bundled together and wrapped in a connective tissue covering. Each muscle is surrounded by a connective tissue sheath called the epimysium. Fascia, connective tissue outside the epimysium, surrounds and separates the muscles. Portions of the epimysium project inward to divide the muscle into compartments. Each compartment contains a bundle of muscle fibers. Each bundle of muscle fiber is called a fasciculus and is surrounded by a layer of connective tissue called the perimysium. Within the fasciculus, each individual muscle cell, called a muscle fiber, is surrounded by connective tissue called the endomysium.

Skeletal muscle cells (fibers), like other body cells, are soft and fragile. The connective tissue covering furnish support and protection for the delicate cells and allow them to withstand the forces of contraction. The coverings also provide pathways for the passage of blood vessels and nerves.

Commonly, the epimysium, perimysium, and endomysium extend beyond the fleshy part of the muscle, the belly or gaster, to form a thick ropelike tendon or a broad, flat sheet-like aponeurosis. The tendon and aponeurosis form indirect attachments from muscles to the periosteum of bones or to the connective tissue of other muscles. Typically a muscle spans a joint and is attached to bones by tendons at both ends. One of the bones remains relatively fixed or stable while the other end moves as a result of muscle contraction.

Skeletal muscles have an abundant supply of blood vessels and nerves. This is directly related to the primary function of skeletal muscle, contraction. Before a skeletal muscle fiber can contract, it has to receive an impulse from a nerve cell. Generally, an artery and at least one vein accompany each nerve that penetrates the epimysium of a skeletal muscle. Branches of the nerve and blood vessels follow the connective tissue components of the muscle of a nerve cell and with one or more minute blood vessels called capillaries.

Muscle Types

In the body, there are three types of muscle: skeletal (striated), smooth, and cardiac.

Skeletal Muscle

Skeletal muscle, attached to bones, is responsible for skeletal movements. The peripheral portion of the central nervous system (CNS) controls the skeletal muscles. Thus, these muscles are under conscious, or voluntary, control. The basic unit is the muscle fiber with many nuclei. These muscle fibers are striated (having transverse streaks) and each acts independently of neighboring muscle fibers.

Smooth Muscle

Smooth muscle, found in the walls of the hollow internal organs such as blood vessels, the gastrointestinal tract, bladder, and uterus, is under control of the autonomic nervous system. Smooth muscle cannot be controlled consciously and thus acts involuntarily. The non-striated (smooth) muscle cell is spindle-shaped and has one central nucleus. Smooth muscle contracts slowly and rhythmically.

Cardiac Muscle

Cardiac muscle, found in the walls of the heart, is also under control of the autonomic nervous system. The cardiac muscle cell has one central nucleus, like smooth muscle, but it also is striated, like skeletal muscle. The cardiac muscle cell is rectangular in shape. The contraction of cardiac muscle is involuntary, strong, and rhythmical.

Smooth and cardiac muscle will be discussed in detail with respect to their appropriate systems. This unit mainly covers the skeletal muscular system

Muscle Groups

There are more than 600 muscles in the body, which together account for about 40 percent of a person's weight.

Most skeletal muscles have names that describe some feature of the muscle. Often several criteria are combined into one name. Associating the muscle's characteristics with its name will help you learn and remember them. The following are some terms relating to muscle features that are used in naming muscles.

• Size: vastus (huge); maximus (large); longus (long); minimus (small); brevis (short).
• Shape: deltoid (triangular); rhomboid (like a rhombus with equal and parallel sides); latissimus (wide); teres (round); trapezius (like a trapezoid, a four-sided figure with two sides parallel).
• Direction of fibers: rectus (straight); transverse (across); oblique (diagonally); orbicularis (circular).
• Location: pectoralis (chest); gluteus (buttock or rump); brachii (arm); supra- (above); infra- (below); sub- (under or beneath); lateralis (lateral).
• Number of origins: biceps (two heads); triceps (three heads); quadriceps (four heads).
• Origin and insertion: sternocleidomastoideus (origin on the sternum and clavicle, insertion on the mastoid process); brachioradialis (origin on the brachium or arm, insertion on the radius).
• Action: abductor (to abduct a structure); adductor (to adduct a structure); flexor (to flex a structure); extensor (to extend a structure); levator (to lift or elevate a structure); masseter (a chewer).